Homo in the Middle Pleistocene: Hypodigms, Variation, and Species Recognition

作者: G. Philip Rightmire

DOI: 10.1002/EVAN.20160

关键词: Human evolutionHomo heidelbergensisHomo sapiensPrehistoryTaxonomy (biology)CraniaSystematicsEvolutionary biologyPleistoceneBiology

摘要: It is generally accepted that modern humans evolved in Africa. This consensus has emerged the last two decades, as molecular evidence been coupled with findings from paleontology and prehistory. Patterns of DNA variation living populations, morphology fossils, archeological traces can all be read to show our species deep roots Africa began disperse into other regions only late Pleistocene. There are still questions about timing these dispersals extent which people replaced or exchanged genes other, more archaic groups, but new problems now coming clearly focus. These relate ancestry Homo sapiens Middle In this essay, I emphasize available Pleistocene localities Europe, exploring among individuals, composition hypodigms, species-level taxonomy, evolutionary relationships hominin populations. One obvious difficulty fossils scarce. For most part, incomplete crania African localities. The record comprehensive for Sima de los Huesos Spain, principally jaws known European sites. means skull traits measurements must provide basis sorting individuals groups building differential diagnoses. much material, dating poorly controlled, although a few important assemblages placed securely within chronological framework. Despite constraints, it possible point toward tentative solutions.

参考文章(43)
LS Barham, KA Robson Brown, None, Human Roots*: Africa and Asia in the Middle Pleistocene ,(2001)
Fred Spoor, Paul O'Higgins, Christopher Dean, Daniel E. Lieberman, Anterior sphenoid in modern humans Nature. ,vol. 397, pp. 572- 572 ,(1999) , 10.1038/17505
Daniel E. Lieberman, Sphenoid shortening and the evolution of modern human cranial shape Nature. ,vol. 393, pp. 158- 162 ,(1998) , 10.1038/30227
Juan-Luis Arsuaga, Carlos Lorenzo, José-Miguel Carretero, Ana Gracia, Ignacio Martínez, Nuria García, José María, Bermúdez de Castro, Eudald Carbonell, A complete human pelvis from the Middle Pleistocene of Spain Nature. ,vol. 399, pp. 255- 258 ,(1999) , 10.1038/20430
R.J. Clarke, The Ndutu cranium and the origin ofHomo sapiens Journal of Human Evolution. ,vol. 19, pp. 699- 736 ,(1990) , 10.1016/0047-2484(90)90004-U
Antonio Rosas, José M. Bermúdez De Castro, The Mauer mandible and the evolutionary significance of Homo heidelbergensis Geobios. ,vol. 31, pp. 687- 697 ,(1998) , 10.1016/S0016-6995(98)80055-7
Juan-Luis Arsuaga, Ignacio Martínez, Carlos Lorenzo, Ana Gracia, Alberto Mun˜oz, Oscar Alonso, Jesu´s Gallego, The human cranial remains from Gran Dolina Lower Pleistocene site (Sierra de Atapuerca, Spain). Journal of Human Evolution. ,vol. 37, pp. 431- 457 ,(1999) , 10.1006/JHEV.1999.0309
J. Clark, J de Heinzelin, K. Schick, W. Hart, T. White, G WoldeGabriel, R. Walter, G Suwa, B Asfaw, E Vrba, al. et, African Homo erectus: old radiometric ages and young Oldowan assemblages in the Middle Awash Valley, Ethiopia. Science. ,vol. 264, pp. 1907- 1910 ,(1994) , 10.1126/SCIENCE.8009220