A new family and genus of acalypterate flies from the Neotropical region, with a phylogenetic analysis of Carnoidea family relationships (Diptera, Schizophora)

摘要: Abstract.  The acalypterate family Inbiomyiidae fam.n. (Diptera, Carnoidea) is described for the newly discovered Neotropical genus Inbiomyiagen.n. with its type species I. mcalpineorumsp.n. from Costa Rica. ranges Guatemala south to French Guiana and Bolivia includes a total of fourteen undescribed species, ten which will be formally in separate paper. Inbiomyia distinctive, characteristic, extremely shortened head nonfunctional ptilinum reduced chaetotaxy, first flagellomere very elongate, dorsoapically inserted arista, proboscis largely labellar lobes that point different directions, mid tibia lacking apicoventral bristle, unusual fusion male sternites 5–7, sternite 8, elongate surstyluslike ventral epandrial lobes, cerci absent both sexes, truncate female genitalia, large, flattened eggs. larva biology are unknown. occurs mostly primary lowland rain forest often associated decaying foliage fallen trees. belong superfamily Carnoidea. previously doubtful monophyly Carnoidea accepted tentatively on basis established synapomorphies genitalia. Family level relationships analysed quantitatively time based matrix fifty-eight morphological characters. putative sister group relationship monotypic Australasian Australimyzidae supported by several synapomorphies, postabdomen. status confirmed, rejecting previous claims (or synonymy) Carnidae. analysis also leads revised hypotheses Cryptochetidae Acartophthalmidae, paraphyly ‘Tethinidae’ regard Canacidae, suspected authors, confirmed.