摘要: Publisher Summary Central pattern generator circuits provide rhythmic patterns of activity among connected neurons that mediate walking. Local perturbations, such as irregularities in ground height, are relayed to central generators by feedback from many thousands mechanoreceptors. This sensory information is integrated locally spiking and nonspiking local interneurons control maintain patterned firing motoneurons. integrate about leg position, allowing compensatory adjustments at the joints for maintaining posture gait. In insect species, afferents thoracic nervous system could alone forward locomotion. Circuits underlying locomotion toward a stimulus source appear be more complexly organized than those mediating escape behaviors. Descending relay brain ganglia have several discrete dendritic domains brain, which usually receive inputs indirectly through ensembles interneurons. addition, descending supply motor also connections prominent midline neuropil, called “the complex.” chapter discusses some known properties this center speculates significance its structure. To date, behavioral genetics, electrophysiology, comparative anatomy all suggest complex plays cerebellar-like role controlling locomotion, change direction, other limb actions. However, before considering complex, it important outline segmental organization emphasize unique position within three pre-oral ganglia.