摘要: STUDIES of the role haplodiploidy in evolution eusociality have been limited to Hymenoptera, only insects known exhibit both reproductive castes and haplodiploid genetic system1. Because aculeate Hymenoptera share many other traits that may affect sociality, such as provisioning at nests, powerful flight, mandibulate mouthparts, stings, it has difficult separate effects from characteristics this taxonomic group2'3. Here I report presence a second insect taxon, order Thysanoptera. Sub-fertile 'soldier' adults Australian gall thrips Oncothrips tepperi Karny O. habrus Mound defend containing their mother siblings invasion takeover by inquiline species invaders. provide remarkable new opportunities for analysing causes eusociality.