Comparative floral structure and systematics in Crossosomatales (Crossosomataceae, Stachyuraceae, Staphyleaceae, Aphloiaceae, Geissolomataceae, Ixerbaceae, Strasburgeriaceae)
作者： MERRAN L. MATTHEWS , PETER K. ENDRESS
DOI: 10.1111/J.1095-8339.2005.00347.X
关键词: Petal 、 Stamen 、 Anatomy 、 Locule 、 Gynoecium 、 Strasburgeriaceae 、 Botany 、 Biology 、 Perianth 、 Staphyleaceae 、 Hypanthium
摘要: Floral structure of all putative families Crossosomatales as suggested by molecular studies was comparatively studied. The seven comprise Crossosomataceae, Stachyuraceae, Staphyleaceae, Aphloiaceae, Geissolomataceae, Ixerbaceae, and Strasburgeriaceae. entire clade (1) is highly supported floral structure, also the clades (in sequence diminishing structural support): Ixerbaceae/Strasburgeriaceae (2), Geissolomataceae/Ixerbaceae/Strasburgeriaceae (3), Aphloiaceae/Geissolomataceae/Ixerbaceae/Strasburgeriaceae (4), Crossosomataceae/Stachyuraceae/Staphyleaceae (5). Among prominent features are solitary flowers, presence a cup, imbricate sepals with outermost smaller than inner, pollen grains horizontally extended endoapertures, shortly stalked gynoecium, postgenitally united carpel tips forming compitum, stigmatic papillae two- or more-cellular, ovary locules tapering upwards, long integuments zigzag micropyles, cell clusters bundles yellow crystals, mucilage cells, seeds smooth, sclerified testa without differentiated tegmen. Clade (2) characterized large petals tight, pointed cone in bud, stamens long, stout filaments sagittate anthers, streamlined, conical antitropous ovules, rudimentary aril, lignified, unicellular, T-shaped hairs idioblasts striate mucilaginous walls. (3) alternisepalous carpels, punctiform stigma formed twisted tips, synascidiate ovary, only one two pendant ovules per carpel, nectary recesses between androecium gynoecium. (4) pronounced ‘pollen buds’. (5) polygamous functionally unisexual x-shaped free follicular carpels (not Stachyuraceae). Crossosomataceae although not retrieved studies, share several special features: polystemonous androecium; basifixed anthers connective protrusion; more less decurrent crests; camplyotropous reniform seeds; simple, disc-shaped nectaries absence hairs. © 2005 Linnean Society London, Botanical Journal Society, 2005, 147, 1–46.
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